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Honestly I think you should remove the body filler and try to knock out as much of the dent as possible. I don't think it'll take very long on the fenders. You will never get get them completely flat again because the metal is streched, unless u have a press and mold. A half inch of bondo is leaning on the overly thick side of things and will eventualy fail. I suggest taking the time to do a better job on it and not having to do it again. Use a small hammer with a flat and blemish free surface and a dollie and just beat it as close as you can get it. Then fill what is left. Make sure you sand the area to be filled with some 40 grit and then clean it with paint thinner. Then use the bondo.
1/4" is to thick for body filler. I never prime under filler either.