For what year model truck are you seeking an answer?For a 1983 C10 with MV4 (350C) transmission, the orange test connector is used to monitor and verify when 12-volt accessory power is supplied to the transmission TCC circuit through the service brake [applied] switch and low engine vacuum (i.e., high engine load) switch. There is also a minimum road speed limit switch inside the transmission. All of the described switches are connected in series with the TCC solenoid. See diagram.
You're reading it wrong. Key on 12v to the solenoid. The brake switch or low vacuum switch can interrupt the circuit open. The pressure switch in the valve body grounds the solenoid when the appropriate pressure is met and applies the torque converter clutch. If you have a test light bulb installed on the test connector you can hit the go pedal and see it go out, let off see it come on, hit the brakes, see it go out, let off the brakes, on and so forth.